What follows is a list of Mesozoic maniraptor genera. I include some information on each taxon, including the year they were named and where they were found. If you see any errors or omissions, please leave a comment!
For Cenozoic extinct birds (including some with possible Cretaceous records), see this list.
Resources I've found particularly helpful in compiling this list include Holtz's Winter 2011 Dinosaur Genus List, The Compact Thescelosaurus, The Theropod Database, DinoData (now defunct), and A Field Guide to Mesozoic Birds and Other Winged Dinosaurs by Matt Martyniuk.
Alvarezsaurs
-Achillesaurus (2007): Late Cretaceous (Santonian) Argentina. A large alvarezsaurid. May have been the same as Alvarezsaurus.
-Albertonykus (2009): Late Cretaceous (Maastrichtian) Canada. The first North American alvarezsaurid to be named. Found in an Albertosaurus bonebed. May have fed on wood-eating termitoids.
-Albinykus (2011): Late Cretaceous (Santonian–Campanian) Mongolia. Was discovered in a sitting position.
-Alnashetri (2012): Late Cretaceous (Cenomanian–Turonian) Argentina. Known from hindlimbs. May have been some other type of coelurosaur.
-Alvarezsaurus (1991): Late Cretaceous (Santonian) Argentina. Known from a partial skeleton.
-Bannykus (2018): Early Cretaceous (Aptian) China. Had forelimbs intermediate in length between those of Haplocheirus and parvicursorines. The second finger might have been used in probing for food in crevices.
-Bonapartenykus (2012): Late Cretaceous (Campanian–Maastrichtian) Argentina. The first alvarezsaur found with associated eggs. Closely related to Patagonykus. The largest known alvarezsaur, at two and a half meters long.
-Ceratonykus (2009): Late Cretaceous (Campanian?–Maastrichtian?) Mongolia. Seemingly had strange spurs on its hands. Possibly the same as Parvicursor.
-Dzharaonyx (2022): Late Cretaceous (Turonian) Uzbekistan. The oldest known parvicursorine.
-Haplocheirus (2010): Late Jurassic (Oxfordian) China. One of the earliest known alvarezsaurs. It retained numerous ancestral features, including hands with three functional fingers (with a large thumb claw) and several large teeth. The size and shape of its sclerotic ring suggest it may have been nocturnal.
-Heptasteornis (1975): Late Cretaceous (Maastrichtian) Romania. Initially thought to have been an owl.
-Jaculinykus (2023): Late Cretaceous (Campanian?–Maastrichtian?) Mongolia. One of the most completely known alvarezsaurids. Had two fingers on each hand. The holotype was preserved in a sleeping position.
-Khulsanurus (2021): Late Cretaceous (Campanian?–Maastrichtian?) Mongolia. Known from a partial skeleton.
-Kol (2009): Late Cretaceous (Campanian) Mongolia. A large alvarezsaurid. Its name means "foot" in Mongolian, based on the fact that it is only known from a foot. May have been some other type of maniraptor.
-Linhenykus (2011): Late Cretaceous (Campanian) China. Had only one finger on each hand. (Most other alvarezsaurids still had the vestiges of the second and third fingers.)
-Mononykus (1993): Late Cretaceous (Maastrichtian) Mongolia. First discovered in the 1920s, but was not described until the 1990s. The first alvarezsaurid known from fairly complete remains. Initially thought to have been an avialan. Was initially named "Mononychus", but that name had already been used for a beetle.
-Nemegtonykus (2019): Late Cretaceous (Maastrichtian) Mongolia. Known from a partial skeleton.
-Ondogurvel (2022): Late Cretaceous (Campanian?–Maastrichtian?) Mongolia. Known from a partial skeleton. Had extensively fused foot bones.
-Parvicursor (1996): Late Cretaceous (Campanian?–Maastrichtian?) Mongolia. Known from a small juvenile specimen.
-Patagonykus (1997): Late Cretaceous (Turonian–Coniacian) Argentina. A transitional form between early alvarezsaurids and parvicursorines.
-Qiupanykus (2018): Late Cretaceous (Maastrichtian?) China. Known from a partial skeleton.
-Shishugounykus (2019): Late Jurassic (Oxfordian) China. An early alvarezsaur known from a partial skeleton.
-Shuvuuia (1998): Late Cretaceous (Campanian) Mongolia. Known from several excellent fossils. The first alvarezsaurid found with preserved feathers. The size and shape of its sclerotic ring suggest it may have been nocturnal, and its inner ear anatomy suggests that it had very sensitive hearing.
-Trierarchuncus (2020): Late Cretaceous (Maastrichtian) U.S.A. Thumb claws of several different growth stages are known.
-Xixianykus (2010): Late Cretaceous (Santonian) China. Had very long legs in relation to body size and so was probably very fast even for an alvarezsaur.
-Xiyunykus (2018): Early Cretaceous (Barremian–Aptian) China. An early alvarezsaur with incipient adaptations for increased power of the forelimbs (possibly for use in digging).
Therizinosaurs
-Alxasaurus (1993): Early Cretaceous (Aptian) Mongolia. Its discovery showed that therizinosaurs were maniraptors. Might have been a more generalist feeder than other therizinosaurs.
-Beipiaosaurus (1999): Early Cretaceous (Barremian–Aptian) China. The first therizinosaur found with preserved feathers. EBFFs (Elongated Broad Filamentous Feathers) were first discovered in this taxon. Had a pygostyle that had evolved independently of pygostylians.
-Enigmosaurus (1983): Late Cretaceous (Cenomanian–Coniacian) Mongolia. Known from only a pelvis. Probably the same as Erlikosaurus.
-Erliansaurus (2002): Late Cretaceous (Santonian?) China. A transitional form between early therizinosaurs and therizinosaurids.
-Erlikosaurus (1980): Late Cretaceous (Cenomanian–Coniacian) Mongolia. Was found with a well-preserved skull. Biomechanical studies show it had a weak bite force consistent with a herbivorous diet. Therizinosaur eggs found in China may belong to this taxon.
-Eshanosaurus (2001): Early Jurassic (Hettangian) China. Known from a partial jaw. Possibly the oldest known therizinosaur. May have actually been an early sauropodomorph.
-Falcarius (2005): Early Cretaceous (Berriasian–Valanginian) U.S.A. Possessed several ancestral features such as possible omnivory, a forward-pointing pubis, and long legs with three weight-bearing toes on each foot. Known from a mass accumulation of dozens or hundreds of individuals.
-Jianchangosaurus (2013): Early Cretaceous (Aptian) China. An early therizinosaur known from a fairly complete juvenile skeleton. Possibly a younger individual of Beipiaosaurus.
-Lingyuanosaurus (2019): Early Cretaceous (Aptian) China. Known from a partial skeleton, possibly of a juvenile.
-Martharaptor (2012): Early Cretaceous (Berriasian–Valanginian) U.S.A. Known from some limb bones, vertebrae fragments, and a few other parts. May have been some other type of maniraptor.
-Paralitherizinosaurus (2022): Late Cretaceous (Campanian) Japan. Known from a partial vertebra and hand.
-Nanshiungosaurus (1979): Late Cretaceous (Maastrichtian) China. Initially described as a sauropod.
-Neimongosaurus (2001): Late Cretaceous (Santonian?) China. Had a long neck and a deep jaw.
-Nothronychus (2001): Late Cretaceous (Turonian) U.S.A. The first therizinosaur known from North America. Two species have been described, N. mckinleyi and N. graffami.
-Segnosaurus (1979): Late Cretaceous (Cenomanian–Coniacian) Mongolia. Initially thought to have been a fish-eating theropod.
-Suzhousaurus (2007): Early Cretaceous (Aptian–Albian) China. May have been closely related to Nothronychus.
-Therizinosaurus (1954): Late Cretaceous (Maastrichtian) Mongolia. Initially thought to have been a giant turtle-like reptile. The largest known therizinosaur, possibly nearing ten meters long and more than three meters tall at the hip. Known from only limb bones, including its large powerful arms with long claws. Each claw may have been almost a meter long in life.
Oviraptorosaurs
-Anomalipes (2018): Late Cretaceous (Campanian?) China. Known from a partial hindlimb.
-Anzu (2014): Late Cretaceous (Maastrichtian) U.S.A. The largest known oviraptorosaur in North America and the most completely known caenagnathid. Had a pygostyle that had evolved independently of pygostylians.
-Apatoraptor (2016): Late Cretaceous (Campanian) Canada. Known from a partial skeleton. Quill knobs have been identified on its ulna.
-Avimimus (1981): Late Cretaceous (Campanian–Maastrichtian) China and Mongolia. Had a strange combination of features such as a long neck, a short tail, long legs, and fused palm bones. The first non-avialan dinosaur found with evidence of feathers, in the form of a ridge on the ulna that may have been an attachment point for feathers. Trackways and at least one bonebed suggest that it may have been a gregarious animal. Two species have been named, A. portentosus and A. nemegtensis.
-Banji (2010): Late Cretaceous (Maastrichtian) China. Known from a subadult skull. Had strange ridges on its crest, which may indicate a keratinous sheath. May have been more carnivorous compared to contemporary oviraptorosaurs.
-Beibeilong (2017): Late Cretaceous (Cenomanian–Turonian) China. Known from a large embryo and eggs.
-Caenagnathasia (1994): Late Cretaceous (Turonian–Santonian?) China and Uzbekistan. Known from toothless jaws and other fragments.
-Caenagnathus (1940): Late Cretaceous (Campanian) Canada. A large caenagnathid. Once thought to be the same as Chirostenotes, but some remains do appear to be from a distinct animal.
-Calamospondylus (1866): Early Cretaceous (Barremian) U.K. Known from only isolated vertebrae.
-Caudipteryx (1998): Early Cretaceous (Barremian–Aptian) China. Very common in one specific region of the Yixian Formation. Had only two functional fingers. Its fossilized feathers appear to show a barred pattern on its tail frond. Has been found with gizzard stones. Two species have been named, C. zoui and C. dongi. C. zoui appears to lack secondary feathers, and analysis of the melanosomes preserved in its feathers indicates that it may have had black body feathers.
-Chirostenotes (1924): Late Cretaceous (Campanian) Canada. The first oviraptorosaur known from North America. It has been suggested that it used its second finger for probing crevices in search of food, but if it had primary feathers like other pennaraptors this would not have been feasible. Includes "Macrophalangia".
-Citipati (2001): Late Cretaceous (Campanian) Mongolia. Known from several complete skeletons, including individuals brooding their nests. Had a pygostyle that had evolved independently of pygostylians. Often mislabeled "Oviraptor" in many restorations.
-Citipes (2020): Late Cretaceous (Campanian) Canada. Once thought to be a species of Chirostenotes or Leptorhynchos. Remains of juveniles have been found as gut contents of juvenile Gorgosaurus.
-Conchoraptor (1986): Late Cretaceous (Maastrichtian) Mongolia. Had a very small crest and a pygostyle that had evolved independently of pygostylians. Its name means "shellfish thief", based on the idea that it may have fed on the small bivalves that were found in the same deposits.
-Corythoraptor (2017): Late Cretaceous (Maastrichtian) China. Had a very large crest, likely superficially similar to that of cassowaries. Possibly an older individual of Banji.
-Elmisaurus (1981): Late Cretaceous (Maastrichtian) Mongolia. Closely related to Chirostenotes. The first oviraptorosaur found to have had a pygostyle that had evolved independently of pygostylians. Includes "Nomingia".
-Eoneophron (2024): Late Cretaceous (Maastrichtian) U.S.A. Known from a partial hindlimb.
-Epichirostenotes (2011): Late Cretaceous (Campanian) Canada. The holotype was once thought to be a specimen of Chirostenotes.
-Ganzhousaurus (2013): Late Cretaceous (Maastrichtian) China. Known from a partial skeleton. May have been more herbivorous compared to contemporary oviraptorosaurs.
-Gigantoraptor (2007): Late Cretaceous (Santonian?) China. The largest known oviraptorosaur, being almost nine meters long. Had the longest legs of any theropod. Giant theropod nests found in China and Mongolia may belong to this taxon. It grew faster than similarly-sized tyrannosaurids did.
-Gobiraptor (2019): Late Cretaceous (Maastrichtian) Mongolia. Known from a juvenile specimen. May have been the same as Conchoraptor.
-Hagryphus (2005): Late Cretaceous (Campanian) U.S.A. A large oviraptorosaur known from forelimb bones.
-Heyuannia (2002): Late Cretaceous (Campanian?–Maastrichtian) China and Mongolia. Known from several excellent fossils. Chemical analysis suggests that its eggs were blue-green in color. Two species have been named, H. huangi and H. yanshini. H. huangi probably had only two functional fingers. H. yanshini was initially named "Ingenia" in 1981, but that name had already been used for a nematode.
-Huanansaurus (2015): Late Cretaceous (Maastrichtian) China. Known from a partial skeleton with a nearly complete skull. Probably closely related to Citipati.
-Incisivosaurus (2002): Early Cretaceous (Barremian–Aptian) China. Had long front teeth akin to incisors with small peg-shaped teeth behind. Juvenile and half-grown specimens preserved with feathers show possible growth stages and feather development. The youngest specimen lacks secondaries. Possibly the same as Protarchaeopteryx.
-Jiangxisaurus (2013): Late Cretaceous (Maastrichtian) China. Known from a partial skeleton of a non-adult individual. May have been more herbivorous compared to contemporary oviraptorosaurs. Possibly the same as Ganzhousaurus.
-Khaan (2001): Late Cretaceous (Campanian) Mongolia. Known from several complete skeletons. Probably closely related to Conchoraptor.
-Leptorhynchos (2013): Late Cretaceous (Campanian) U.S.A. Known from jaws and possibly some other bones.
-Luoyanggia (2009): Early Cretaceous (Aptian–Albian) China. An early oviraptorosaur known from a jaw, hip, and foot bones.
-Machairasaurus (2010): Late Cretaceous (Campanian–Maastrichtian) China. Known from a partial forelimb and hand. An oviraptorosaur specimen preserved on its nest has been referred to this taxon. Had weakly-curved claws, suggesting herbivory.
-Microvenator (1970): Early Cretaceous (Aptian–Albian) U.S.A. The teeth of Deinonychus were once thought to belong to this taxon.
-Nankangia (2013): Late Cretaceous (Maastrichtian) China. Known from a lower jaw, vertebrae, and limb bones. May have been more herbivorous compared to contemporary oviraptorosaurs.
-Nemegtomaia (2005): Late Cretaceous (Maastrichtian) Mongolia. Was initially named "Nemegtia", but that name had already been used for a crustacean. Had a very tall crest. One known specimen has been preserved on its nest.
-Ningyuansaurus (2012): Early Cretaceous (Aptian) China. Had more teeth than other early oviraptorosaurs. Known to have eaten seeds. May have been some other type of maniraptor.
-Ojoraptorsaurus (2011): Late Cretaceous (Maastrichtian) U.S.A. Known from hip bones.
-Oksoko (2020): Late Cretaceous (Maastrichtian) Mongolia. Known from several specimens, including a group of juveniles preserved together. Had only two functional fingers.
-Oviraptor (1924): Late Cretaceous (Campanian) Mongolia. The holotype was found on a nest of eggs mistakenly thought to have belonged to Protoceratops, giving this taxon a reputation as an "egg thief", which its name translates to. Stomach contents show it ate lizards.
-Protarchaeopteryx (1997): Early Cretaceous (Barremian–Aptian) China. The first oviraptorosaur found with preserved feathers.
-Rinchenia (2004): Late Cretaceous (Maastrichtian) Mongolia. Had a very tall crest. Once thought to be a species of Oviraptor.
-Shanyangosaurus (1996): Late Cretaceous (Maastrichtian) China. Known from a poorly preserved, incomplete skeleton. May have been some other type of maniraptor.
-Shixinggia (2005): Late Cretaceous (Maastrichtian) China. Had strange openings in the leg bones.
-Similicaudipteryx (2008): Early Cretaceous (Aptian) China. Similar to Caudipteryx, as its name suggests. Had a pygostyle that had evolved independently of pygostylians.
-Tongtianlong (2016): Late Cretaceous (Campanian–Maastrichtian) China. Known from a nearly complete specimen.
-Wulatelong (2013): Late Cretaceous (Campanian) China. The second toe of the holotype is preserved in a retracted posture.
-Xingtianosaurus (2019): Early Cretaceous (Aptian) China. An early oviraptorosaur known from a partial skeleton.
-Yuanyanglong (2024): Early Cretaceous (Aptian–Albian) China. Known from a pair of specimens preserved together. Had relatively long legs. Has been found with gizzard stones.
-Yulong (2013): Late Cretaceous (Maastrichtian?) China. Known mostly from juvenile specimens.
Deinonychosaurs*
-Acheroraptor (2013): Late Cretaceous (Maastrichtian) U.S.A. A North American velociraptorine.
-Achillobator (1999): Late Cretaceous (Cenomanian–Coniacian) Mongolia. One of the largest dromaeosaurids. Had very short legs.
-Adasaurus (1983): Late Cretaceous (Maastrichtian) Mongolia. Known from the partial skeleton of an old individual.
-Atrociraptor (2004): Late Cretaceous (Maastrichtian) Canada. Had a very deep snout.
-Austroraptor (2009): Late Cretaceous (Campanian–Maastrichtian) Argentina. A very large unenlagiine with unusually short arms. Its skull was similar in shape to that of a spinosaurid.
-Bambiraptor (2000): Late Cretaceous (Campanian) U.S.A. Known from a half-grown partial skeleton. Might have been able to grasp objects one-handed between the thumb and third finger. May have been a North American microraptorian, or the same as Saurornitholestes. The holotype was once thought to be a North American specimen of Velociraptor.
-Boreonykus (2015): Late Cretaceous (Campanian) Canada. A North American velociraptorine. May have been some other type of coelurosaur.
-Buitreraptor (2005): Late Cretaceous (Cenomanian) Argentina. The most completely known unenlagiine. Had a very long snout and very long arms.
-Changyuraptor (2014): Early Cretaceous (Aptian) China. A fairly large microraptorian. Its tail feathers were the longest known feathers of any Mesozoic dinosaur.
-Dakotaraptor (2015): Late Cretaceous (Maastrichtian) U.S.A. A large dromaeosaurid. Quill knobs have been identified on its ulna. "Wishbones" thought to belong to it actually represent parts of turtle shells, and it has been suspected that some other bones currently assigned to it may belong to other animals as well.
-Daurlong (2022): Early Cretaceous (Aptian) China. Similar to Tianyuraptor and Zhenyuanlong. Known from a nearly complete skeleton with preserved feathers and potential intestinal remnants.
-Deinonychus (1969): Early Cretaceous (Aptian–Albian) U.S.A. The dinosaur that inspired the "Dinosaur Renaissance". Multiple specimens have ben found in association with Tenontosaurus, suggesting that it often fed on this herbivore. This is often considered evidence for group hunting in this taxon, but this is debatable. Juveniles had more recurved claws. A possible egg is known; chemical analysis of it suggests that the eggshell was blue-green in color.
-Dineobellator (2020): Late Cretaceous (Maastrichtian) U.S.A. Its hand and foot claws appear to have been specialized for increased grip strength. Quill knobs have been identified on its ulna.
-Diuqin (2024): Late Cretaceous (Santonian) Argentina. An unenlagiine known from some vertebrae and an arm bone.
-Dromaeosauroides (2003): Early Cretaceous (Berriasian) Denmark. Known only from a tooth.
-Dromaeosaurus (1922): Late Cretaceous (Campanian) Canada and U.S.A. Had very powerful jaws. Was initially thought to have been a small tyrannosauroid.
-Graciliraptor (2004): Early Cretaceous (Barremian–Aptian) China. A microraptorian known from only partial remains.
-Halszkaraptor (2017): Late Cretaceous (Campanian) Mongolia. Likely semi-aquatic. Had numerous teeth, a long neck, and short, stout forelimbs. Unusually among theropods, the third finger was the longest in the hand. May have used its forelimbs for wing-propelled swimming.
-Hesperonychus (2009): Late Cretaceous (Campanian) Canada. The first North American microraptorian to be described (except possibly Bambiraptor). Also the youngest known microraptorian. Known from a pelvis found in 1982, as well as other possible bits and pieces.
-Hulsanpes (1982): Late Cretaceous (Campanian?–Maastrichtian?) Mongolia. Known from only a foot and a partial braincase. Probably closely related to Mahakala and Halszkaraptor.
-Itemirus (1976): Late Cretaceous (Turonian) Uzbekistan. Once thought to have been a tyrannosauroid. Possible remains suggest it may have been comparable in size to Utahraptor.
-Kansaignathus (2021): Late Cretaceous (Santonian) Tajikistan. Known from a lower jaw bone and possibly a few other remains.
-Kuru (2021): Late Cretaceous (Campanian?–Maastrichtian?) Mongolia. Closely related to Velociraptor.
-Linheraptor (2010): Late Cretaceous (Campanian) China. Known from a nearly complete, articulated skeleton. Closely related to and probably the same as Tsaagan.
-Luanchuanraptor (2007): Late Cretaceous (Maastrichtian?) China. The first Chinese dromaeosaurid known from outside of the Gobi Desert and the northeastern region.
-Mahakala (2007): Late Cretaceous (Campanian) Mongolia. A very small, flightless deinonychosaur. Known from a partial specimen discovered in 1992. May have been semi-aquatic, similar to the closely related Halszkaraptor.
-Microraptor (2000): Early Cretaceous (Aptian) China. A small, common microraptorian. Had very long foot feathers, suggesting that it used a method of flight unknown in any modern animal. Thought to have had a feathered crest on its head, but this is likely a taphonomic artifact. The back half of a specimen of this taxon was once combined with the front half of a specimen of Yanornis to create the infamous "Archaeoraptor" hoax. Stomach contents show that it ate enantiornithean avialans, fish, lizards, and mammals. Likely exhibited sequential molt of its wing feathers. Analysis of melanosomes preserved in its feathers shows that it may have had iridescent plumage. The size and shape of its sclerotic ring suggest it may have been nocturnal, but the fact that few nocturnal birds are known to have iridescent plumage casts doubt on this. At least one specimen preserves a pair of long tail feathers at the tip of its tail frond. Includes "Cryptovolans".
-Natovenator (2022): Late Cretaceous (Campanian?–Maastrichtian?) Mongolia. A halszkaraptorine known from a well-preserved partial skeleton, including a nearly complete skull.
-Neuquenraptor (2005): Late Cretaceous (Turonian–Coniacian) Argentina. Probably the same as Unenlagia.
-Ornithodesmus (1887): Early Cretaceous (Barremian) U.K. Known only from hip vertebrae. Initially thought to have been a pterosaur.
-Pamparaptor (2011): Late Cretaceous (Turonian–Coniacian) Argentina. Once thought to represent a juvenile specimen of Neuquenraptor. Known from a troodont-like foot, but was possibly an unenlagiine.
-Pyroraptor (2000): Late Cretaceous (Campanian–Maastrichtian) France. Its name means "fire thief", based on the fact that its remains were found after a forest fire. Probably the same as Variraptor.
-Saurornitholestes (1978): Late Cretaceous (Campanian) Canada and U.S.A. A tooth embedded in a pterosaur arm bone and feeding traces on an ornithomimid tail bone show that it fed on (presumably already dead) azhdarchid pterosaurs and ornithomimids. A partial specimen with tooth marks suggests that it may have been eaten by tyrannosaurids.
-Shanag (2007): Early Cretaceous (Hauterivian–Barremian) Mongolia. A very small dromaeosaurid. May have been a microraptorian or a northern unenlagiine.
-Shri (2021): Late Cretaceous (Campanian?–Maastrichtian?) Mongolia. Known from well-preserved partial skeletons. Closely related to Velociraptor.
-Sinornithosaurus (1999): Early Cretaceous (Barremian–Aptian) China. The first dromaeosaurid found with preserved feathers. The size and shape of its sclerotic ring suggest it may have been cathemeral. Remains have been found as gut contents of Sinocalliopteryx.
-Tianyuraptor (2010): Early Cretaceous (Barremian–Aptian) China. A large, short-armed possible microraptorian.
-Tsaagan (2006): Late Cretaceous (Campanian) Mongolia. Known from a well-preserved skull and some neck vertebrae. Had a strong snout for a velociraptorine.
-Unenlagia (1997): Late Cretaceous (Turonian–Coniacian) Argentina. May have had a greater range of mobility at the shoulder joint than other deinonychosaurs. Two species have been named, U. comahuensis and U. paynemili.
-Utahraptor (1993): Early Cretaceous (Berriasian–Valanginian) U.S.A. The largest known dromaeosaurid, at around seven meters long.
-Variraptor (1998): Late Cretaceous (Campanian–Maastrichtian) France. Known only from very fragmentary and possibly chimeric material.
-Vectiraptor (2021): Early Cretaceous (Barremian) U.K. Known only from partial vertebrae and possibly teeth.
-Velociraptor (1924): Late Cretaceous (Campanian) China and Mongolia. Known from many excellent fossils, including one locked in combat with a Protoceratops. Feeding traces and gut contents show that it scavenged on the remains of Protoceratops and azhdarchid pterosaurs. Quill knobs have been identified on its ulna. Juveniles had more recurved claws. The size and shape of its sclerotic ring suggest it may have been nocturnal. Two species have been named, V. mongoliensis and V. osmolskae.
-Wulong (2020): Early Cretaceous (Aptian) China. A microraptorian with a very long tail, as well as a pair of elongated tail feathers. Analysis of melanosomes preserved in its feathers shows that it may have had iridescent and gray plumage.
-Ypupiara (2021): Late Cretaceous (Maastrichtian) Brazil. An unenlagiine known from jaw fragments. The holotype has been lost in a museum fire.
-Yurgovuchia (2012): Early Cretaceous (Berriasian–Valanginian) U.S.A. Similar in some ways to Utahraptor, but much smaller. Had a fairly flexible tail.
-Zapsalis (1876): Late Cretaceous (Campanian) U.S.A. Known only from teeth. Teeth from numerous locales have been referred to this taxon, but it is unclear whether they all represent the same genus.
-Zhenyuanlong (2015): Early Cretaceous (Aptian) China. Known to have had large wings despite its short arms.
-Zhongjianosaurus (2017): Early Cretaceous (Aptian) China. A very small microraptorian.
*Halszkaraptorines and unenlagiines may have been avialans.
Troodonts**
-Albertavenator (2017): Late Cretaceous (Maastrichtian) Canada. Known from skull fragments.
-Almas (2017): Late Cretaaceous (Campanian) Mongolia. A small troodont known from a partial skeleton, including a nearly complete skull.
-Archaeornithoides (1992): Late Cretaceous (Campanian) Mongolia. Known from a juvenile skull that appears to have been fed on by a deltatheridiid mammal. Once thought to represent a hatchling Tarbosaurus.
-Borogovia (1987): Late Cretaceous (Maastrichtian) Mongolia. Had highly reduced killing claws.
-Byronosaurus (2000): Late Cretaceous (Campanian) Mongolia. Known from a partial skeleton and an incomplete skull.
-Daliansaurus (2017): Early Cretaceous (Barremian–Aptian) China. Known from a nearly complete skeleton. Had a large claw on the fourth toe, similar in size to its killing claw.
-Geminiraptor (2010): Early Cretaceous (Berriasian–Valanginian) U.S.A. The first North American Early Cretaceous troodont described.
-Gobivenator (2014): Late Cretaceous (Campanian) Mongolia. Known from a nearly complete specimen.
-Harenadraco (2024): Late Cretaceous (Campanian?–Maastrichtian?) Mongolia. Had very long, slender feet.
-Hesperornithoides (2019): Late Jurassic (Kimmeridgian–Tithonian) U.S.A. One of the oldest known troodonts. Known from a partial skeleton.
-Hypnovenator (2024): Early Cretaceous (Albian) Japan. Had some similarities to Late Cretaceous troodonts. The holotype was preserved in a sitting position.
-Jianianhualong (2017): Early Cretaceous (Aptian) China. The first troodont found with definite evidence of asymmetric feathers.
-Jinfengopteryx (2005): Early Cretaceous (Aptian) China. Known to have eaten seeds. May have been some other type of paravian.
-Koparion (1994): Late Jurassic (Kimmeridgian) U.S.A. Known only from teeth.
-Liaoningvenator (2017): Early Cretaceous (Barremian–Aptian) China. Known from a nearly complete skeleton.
-Linhevenator (2011): Late Cretaceous (Campanian) China. A derived troodont with extremely short but powerful arms. Had large killing claws for a troodont.
-Mei (2004): Early Cretaceous (Barremian–Aptian) China. Its name means "sleeping" in Chinese, based on the fact that the holotype is a complete skeleton preserved in a sleeping position.
-Papiliovenator (2021): Late Cretaceous (Campanian) China. Known from a partial skeleton, including a nearly complete skull. Its name means "butterfly hunter", referencing the shape of its back vertebrae.
-Pectinodon (1982): Late Cretaceous (Maastrichtian) U.S.A. Known mostly from teeth. Once thought to be the same as Troodon.
-Philovenator (2012): Late Cretaceous (Campanian) China. Had very long, slender feet. Once thought to represent a juvenile specimen of Saurornithoides.
-Saurornithoides (1924): Late Cretaceous (Campanian) Mongolia. Closely related to Troodon.
-Sinornithoides (1994): Early Cretaceous (Aptian–Albian) China. Known from a nearly complete fossil in a sleeping position.
-Sinovenator (2002): Early Cretaceous (Barremian–Aptian) China. Had a backwards-pointing pubis a la dromaeosaurids, indicating that forward-pointing pubes in later troodonts were reversals. One specimen has been preserved in a sleeping position.
-Sinusonasus (2004): Early Cretaceous (Barremian–Aptian) China. Its name means "curved nose", based on the fact that it had curved snout bones. May have been the same as Sinovenator.
-Stenonychosaurus (1932): Late Cretaceous (Campanian) Canada. One of the largest known troodonts, at over three meters long. Once thought to be the same as Troodon. Includes "Latenivenatrix".
-Talos (2011): Late Cretaceous (Campanian) U.S.A. The holotype shows signs of a healed injury to the left killing claw, supporting the idea that troodonts used the second toe claw as a weapon.
-Tamarro (2021): Late Cretaceous (Maastrichtian) Spain. Known from a partial foot bone.
-Tochisaurus (1991): Late Cretaceous (Maastrichtian) Mongolia. Known from only a foot. Probably had reduced killing claws.
-Troodon (1856): Late Cretaceous (Campanian) U.S.A. Once thought to have been a lizard, pachycephalosaur, or ornithopod. Though many North American troodont specimens have been referred to it, it is definitively known only from teeth.
-Urbacodon (2007): Late Cretaceous (Cenomanian–Santonian?) China and Uzbekistan. Known mainly from teeth and jaws. Its name stands for the Uzbek/Russian/British/American/Canadian joint paleontological expeditions. Two species have been named, U. itemirensis and U. norelli.
-Xixiasaurus (2010): Late Cretaceous (Coniacian–Campanian) China. May have been closely related to Urbacodon.
-Zanabazar (2009): Late Cretaceous (Maastrichtian) Mongolia. Once thought to be a species of Saurornithoides.
**Troodonts may have been deinonychosaurs.
Avebrevicaudans
-Abavornis (1998): Late Cretaceous (Turonian) Uzbekistan. Known only from a partial shoulder bone.
-Aberratiodontus (2004): Early Cretaceous (Aptian) China. Initially thought to have been an enantiornithean, but was probably a yanornithiform and possibly the same as Yanornis.
-Abitusavis (2020): Early Cretaceous (Aptian) China. Closely related to Yanornis. Known to have eaten fish.
-Alamitornis (2009): Late Cretaceous (Campanian–Maastrichtian) Argentina. Possibly closely related to Patagopteryx. May have actually been a lizard.
-Alethoalaornis (2007): Early Cretaceous (Aptian) China. An enantiornithean known from several specimens, but not well studied.
-Alexornis (1976): Late Cretaceous (Campanian) Mexico. A sparrow-sized enantiornithean.
-Ambiortus (1982): Early Cretaceous (Hauterivian–Barremian) Mongolia. A euornithean with a mix of ancestral and derived characteristics.
-Antarcticavis (2019): Late Cretaceous (Maastrichtian) Antarctica. A euornithean known from a partial skeleton.
-Apatornis (1873): Late Cretaceous (Coniacian–Campanian) U.S.A. Once thought to be a species of Ichthyornis, but may have been a neornithean.
-Apsaravis (2001): Late Cretaceous (Campanian) Mongolia. A euornithean known from a nearly complete skeleton.
-Archaeorhynchus (2006): Early Cretaceous (Aptian) China. A euornithean with a pair of elongate feathers in the center of its tail fan and a toothless bill with a rounded lower jaw tip. Had large wing feathers as a juvenile, suggesting early onset of flight abilities during growth. Most known specimens have been found with gizzard stones. One specimen was preserved with lung tissues.
-Archaeornithura (2015): Early Cretaceous (Hauterivian) China. The oldest known euornithean. A hongshanornithid known from excellent specimens.
-Asiahesperornis (1991): Late Cretaceous (Maastrichtian) Kazakhstan. Known only from vertebrae and leg bones.
-Asteriornis (2020): Late Cretaceous (Maastrichtian) Belgium. A small pan-galloanseran with long hindlimbs. A nearly complete skull is known.
-Austinornis (2004): Late Cretaceous (Coniacian–Santonian) U.S.A. Possibly an early galliform.
-Avimaia (2019): Early Cretaceous (Aptian) China. A small enantiornithean. The holotype was preserved with an unlaid egg.
-Avisaurus (1985): Late Cretaceous (Maastrichtian) U.S.A. A large, possibly raptorial enantiornithean. Two valid species have been named, A. archibaldi and A. darwini.
-Baptornis (1877): Late Cretaceous (Coniacian–Campanian) Sweden and U.S.A. A nearly complete skeleton is known. Includes "Parascaniornis".
-Bauxitornis (2010): Late Cretaceous (Santonian) Hungary. Described as a large avisaurid, though possibly some other type of maniraptor.
-Beiguornis (2022): Early Cretaceous (Aptian) China. An enantiornithean known from a partial skeleton.
-Bellulornis (2016): Early Cretaceous (Aptian) China. A euornithean known from a mostly complete skeleton lacking a skull. Was initially named "Bellulia", but that name had already been used for a lepidopteran.
-Bohaiornis (2011): Early Cretaceous (Aptian) China. A large enantiornithean known from good specimens.
-Boluochia (1995): Early Cretaceous (Aptian) China. An enantiornithean initially thought to have had a hooked beak, but this was a misinterpretation and it is now known to have had large, curved teeth instead.
-Brevidentavis (2021): Early Cretaceous (Aptian) China. A euornithean known from a partial skull and some vertebrae. Had short, blunt teeth.
-Brevirostruavis (2021): Early Cretaceous (Aptian) China. An enantiornithean with a long hyoid for a non-neornithean theropod.
-Brodavis (2012): Late Cretaceous (Campanian–Maastrichtian) Canada, Mongolia, and U.S.A. A hesperornithean known from freshwater deposits. May have been volant. Probably stayed near the water surface. Four species have been named, B. americanus, B. baileyi, B. mongoliensis, and B. varneri, the last of which was once considered a species of Baptornis.
-Canadaga (1999): Late Cretaceous (Campanian–Maastrichtian) Canada. The youngest and largest known hesperornithean.
-Castignovolucris (2023): Late Cretaceous (Campanian?) France. A large enantiornithean known from a shoulder bone.
-Catenoleimus (1998): Late Cretaceous (Turonian) Uzbekistan. Known only from a badly-preserved shoulder bone.
-Cathayornis (1992): Early Cretaceous (Aptian) China. Once thought to be the same as Sinornis.
-Ceramornis (1963): Late Cretaceous (Maastrichtian) U.S.A. A possible neornithean known from a partial shoulder bone.
-Cerebavis (2006): Late Cretaceous (Cenomanian) Russia. A euornithean known only from a partial skull, once interpreted as a preserved brain.
-Changchengornis (1999): Early Cretaceous (Barremian–Aptian) China. A confuciusornithiform known to have had a feathered crest and a hooked bill.
-Changmaornis (2013): Early Cretaceous (Aptian) China. A euornithean known from leg and hip bones.
-Changzuiornis (2016): Early Cretaceous (Aptian) China. A long-snouted euornithean. Analysis of the melanosomes preserved in its feathers indicates that it may have had black wing and tail feathers.
-Chaoyangia (1993): Early Cretaceous (Aptian) China. Some remains of this taxon have been reassigned to Songlingornis.
-Chiappeavis (2015): Early Cretaceous (Aptian) China. An enantiornithean with a fan-shaped array of tail feathers, superficially similar to that of euornitheans.
-Chongmingia (2016): Early Cretaceous (Aptian) China. Known from a partial skeleton. Probably closely related to Jinguofortis.
-Chupkaornis (2017): Late Cretaceous (Coniacian–Santonian) Japan. Known from vertebrae and parts of the hindlimbs. The oldest and most completely known hesperornithean from Asia.
-Cimolopteryx (1892): Late Cretaceous (Maastrichtian) U.S.A. A possible neornithean known from a shoulder bone.
-Concornis (1992): Early Cretaceous (Barremian) Spain. May have been a wading enantiornithean. May have used bounding and continuous flapping flight.
-Confuciusornis (1995): Early Cretaceous (Aptian) China. The most common Mesozoic dinosaur yet found, known from hundreds or thousands of specimens. Some individuals have been found with a pair of ribbon-shaped tail feathers preserved while others lack them, representing sexual dimorphism. Melanosome distribution suggests it had spotted plumage on the wings, head, and throat. The size and shape of its sclerotic ring suggest it may have been diurnal. May have fed by watching for insects from a perch before making short flights to capture them. Remains have been found as gut contents of Sinocalliopteryx. Three valid species have been named, C. sanctus, C. dui, and C. shifan. Includes "Jinzhouornis".
-Cratoavis (2015): Early Cretaceous (Aptian) Brazil. An enantiornithean known from a small, well-preserved specimen with very long, striped tail feathers.
-Cratonavis (2023): Early Cretaceous (Aptian) China. Probably closely related to Jinguofortis.
-Cruralispennia (2017): Early Cretaceous (Hauterivian) China. An enantiornithean with an unusually short pygostyle. Had strange wire-shaped feathers on its legs and wings, though these may represent juvenile plumage. Grew rapidly for an enantiornithean, nearing adult size within a year.
-Cuspirostrisornis (1997): Early Cretaceous (Aptian) China. An enantiornithean with very large talons.
-Dalingheornis (2006): Early Cretaceous (Aptian) China. An enantiornithean reported to have had heterodactyl feet that had evolved independently of trogons, but this may be an artifact of preservation. Had a longer tail than other enantiornitheans did, though this was probably a juvenile characteristic. Possibly the juvenile of another known genus.
-Dapingfangornis (2006): Early Cretaceous (Aptian) China. An enantiornithean known from a nearly complete skeleton with preserved feathers. Was described as having a horn-like projection on its snout, but this is more likely an artifact of preservation.
-Dingavis (2016): Early Cretaceous (Aptian) China. A long-snouted, toothless euornithean.
-Dunhuangia (2015) Early Cretaceous (Aptian) China. An enantiornithean known from a partial skeleton.
-Elbretornis (2009): Late Cretaceous (Maastrichtian) Argentina. An enantiornithean with very hollow arm bones. May have been the same as Lectavis or Yungavolucris.
-Elektorornis (2019): Late Cretaceous (Cenomanian) Myanmar. An enantiornithean known from a partial specimen preserved in amber. Had an unusually long third toe. One of the smallest known Mesozoic dinosaurs.
-Elsornis (2007): Late Cretaceous (Campanian) Mongolia. A possibly flightless or near-flightless enantiornithean.
-Enaliornis (1876): Early Cretaceous (Albian) U.K. Probably an early, flying hesperornithean. Three species have been named, E. barretti, E. sedgwicki, and E. seeleyi.
-Enantiophoenix (2008): Late Cretaceous (Cenomanian) Lebanon. One of the first dinosaur fossils found in Lebanon. Amber globules associated with the holotype have been interpreted as evidence that it ate tree sap, but this is uncertain given the disarticulated nature of the specimen.
-Enantiornis (1981): Late Cretaceous (Campanian) Argentina. A large enantiornithean.
-Eoalulavis (1996): Early Cretaceous (Barremian) Spain. The oldest known maniraptor with an alula at the time of its discovery. Stomach contents show that it may have eaten aquatic crustaceans. May have used bounding and continuous flapping flight.
-Eocathayornis (2002): Early Cretaceous (Aptian) China. In spite of its name, it may not be closely related to Cathayornis.
-Eoconfuciusornis (2008): Early Cretaceous (Hauterivian) China. An early confuciusornithiform. Analysis of the melanosomes preserved in its feathers indicates that it may have been mostly black, with a gray and iridescent head, gray legs, and a brown throat patch.
-Eoenantiornis (1999): Early Cretaceous (Barremian–Aptian) China. A small enantiornithean with a short, blunt snout.
-Eogranivora (2018): Early Cretaceous (Barremian–Aptian) China. A toothless, seed eating euornithean. It did not have a first toe, suggesting that it was highly terrestrial. The holotype was once thought to be a specimen of Hongshanornis.
-Eopengornis (2014): Early Cretaceous (Hauterivian) China. An enantiornithean with a pair of fully pennaceous tail feathers, suggesting that the ribbon-like morphology found in the tail feathers of other enantiornitheans may have been modified from fully pennaceous feathers.
-Evgenavis (2014): Early Cretaceous (Barremian–Aptian) Russia. Known from a tarsometatarsus. Had features similar to both ornithothoraceans and confuciusornithiforms.
-Explorornis (1998): Late Cretaceous (Turonian) Uzbekistan. An enantiornithean known only from a partial shoulder bone.
-Falcatakely (2020): Late Cretaceous (Maastrichtian) Madagascar. An enantiornithean with a superficially toucan-like skull.
-Feitianius (2015): Early Cretaceous (Aptian) China. An enantiornithean with ornamental tail plumage formed from three feather morphotypes.
-Flexomornis (2010): Late Cretaceous (Cenomanian) U.S.A. An enantiornithean that was found in marine deposits.
-Fortipesavis (2021): Late Cretaceous (Cenomanian) Myanmar. An enantiornithean known from a partial foot preserved in amber. Had an unusually thick fourth toe.
-Fortunguavis (2014): Early Cretaceous (Aptian) China. An enantiornithean with stout feet and strongly recurved claws. Has been suggested to have been specialized for trunk-climbing, but this has been disputed.
-Fumicollis (2015): Late Cretaceous (Coniacian–Campanian) U.S.A. A small hesperornithean known from a partial skeleton. Gut contents show it ate fish.
-Gallornis (1931): Early Cretaceous (Berriasian–Hauterivian) France. Known only from fragmented limb bones.
-Gansus (1984): Early Cretaceous (Aptian–Albian) China. A foot-propelled diving euornithean. Known from many good skeletons, including some with integument preserved. Plumage was probably dark colored.
-Gargantuavis (1998): Late Cretaceous (Campanian–Maastrichtian) France and Spain. One of the largest Mesozoic avialans. Generally thought to have been a euornithean, but may have been some other type of avialan. Fossil eggs found in France may belong to this taxon.
-Gettyia (2018): Late Cretaceous (Campanian) U.S.A. Probably closely related to Avisaurus, which it was once considered a species of.
-Gobipipus (2013): Late Cretaceous (Campanian?–Maastrichtian?) Mongolia. An enantiornithean known only from embryos.
-Gobipteryx (1976): Late Cretaceous (Campanian–Maastrichtian?) Mongolia. One of the few enantiornitheans known to have had toothless jaws. One specimen was originally thought to have been a species of Nanantius. Possible eggs are known.
-Grabauornis (2014): Early Cretaceous (Aptian) China. An enantiornithean known from a nearly complete specimen.
-Gracilornis (2011): Early Cretaceous (Aptian) China. A slender enantiornithean. Known from a complete skeleton.
-Graculavus (1872): Late Cretaceous (Maastrichtian) U.S.A. A turkey-sized possible neornithean. Two species have been named, G. velox and G. augustus. Includes "Limosavis".
-Gretcheniao (2019): Early Cretaceous (Barremian–Aptian) China. A large enantiornithean with long hands. May have used continuous flapping flight.
-Guildavis (2004): Late Cretaceous (exact age unknown) U.S.A. Once thought to be a species of Ichthyornis.
-Gurilynia (1999): Late Cretaceous (Maastrichtian) Mongolia. A large enantiornithean.
-Halimornis (2002): Late Cretaceous (Santonian–Campanian) U.S.A. An enantiornithean that was found in marine deposits.
-Hesperornis (1872): Late Cretaceous (Coniacian–Campanian) Canada, Russia, Sweden, and U.S.A. An aquatic euornithean known from many excellent fossils from both marine and freshwater deposits. Remains have been found as gut contents of Tylosaurus and one individual possesses healed bite marks made by a polycotylid plesiosaur. Likely a deep-diving taxon. Eleven species have been named, H. regalis, H. altus, H. bairdi, H. chowi, H. crassipes, H. gracilis, H. lumgairi, H. macdonaldi, H. mengeli, H. montanus, and H. rossicus. Includes "Coniornis".
-Holbotia (2015): Early Cretaceous (Hauterivian–Barremian) Mongolia. May have had a more flexible neck than other enantiornitheans.
-Hollanda (2010): Late Cretaceous (Campanian?–Maastrichtian?) Mongolia. Probably a roadrunner-like running ground avialan.
-Hongshanornis (2005): Early Cretaceous (Barremian–Aptian) China. A euornithean known from complete specimens with preserved feathers. May have had a feathered crest on its head. Likely had a wading ecology.
-Horezmavis (1983): Late Cretaceous (Cenomanian) Uzbekistan. Known from only a foot.
-Houornis (2015): Early Cretaceous (Aptian) China. Once thought to be a species of Cathayornis. The holotype is preserved as an impression.
-Huoshanornis (2010): Early Cretaceous (Aptian) China. An enantiornithean that may have been very agile in flight. Known from a nearly complete skeleton.
-Iaceornis (2004): Late Cretaceous (Coniacian–Campanian) U.S.A. Once thought to be a species of Ichthyornis.
-Iberomesornis (1992): Early Cretaceous (Barremian) Spain. A sparrow-sized enantiornithean, one of the smallest known Mesozoic dinosaurs.
-Ichthyornis (1873): Late Cretaceous (Cenomanian–Campanian) Canada, Mexico, and U.S.A. A toothed, gull-like marine euornithean. One of the first fossil avialans found in North America.
-Imparavis (2024): Early Cretaceous (Aptian) China. One of the oldest known toothless enantiornitheans. Had very pronounced muscle attachment points on its forelimb bones.
-Incolornis (1998): Late Cretaceous (Turonian) Uzbekistan. Two species have been named, I. silvae and I. martini, the latter of which was once thought to be a species of Enantiornis.
-Intiornis (2010): Late Cretaceous (Campanian) Argentina. A sparrow-sized enantiornithean well adapted for perching. Known only from a foot.
-Iteravis (2014): Early Cretaceous (Aptian) China. A very common euornithean with ornamental tail feathers. The earliest dinosaur known to have had salt glands. May have had lobed toes. Analysis of the melanosomes preserved in its feathers indicates that it may have had black wing and body feathers.
-Janavis (2022): Late Cretaceous (Maastrichtian) Belgium. Similar to but larger than Ichthyornis.
-Jianchangornis (2009): Early Cretaceous (Aptian) China. A euornithean known from a nearly complete, articulated skeleton, though the skull is a composite.
-Jibeinia (1997): Early Cretaceous (Aptian) China. Superficially similar to confuciusornithiforms, but was probably an enantiornithean.
-Jinguofortis (2018): Early Cretaceous (Barremian) China. Known from a complete specimen with well-preserved wing feathers.
-Jiuquanornis (2013): Early Cretaceous (Aptian) China. A euornithean known from a breastbone, wishbone, and ribs.
-Judinornis (1983): Late Cretaceous (Maastrichtian) Mongolia. A freshwater hesperornithean.
-Juehuaornis (2015): Early Cretaceous (Aptian) China. A euornithean with a long, hook-tipped snout.
-Junornis (2017): Early Cretaceous (Barremian–Aptian) China. An enantiornithean known from a well-preserved specimen. May have used bounding flight.
-Kaririavis (2021): Early Cretaceous (Aptian) Brazil. A small euornithean known from a partial foot. The oldest known euornithean from South America.
-Khinganornis (2020): Early Cretaceous (Aptian) China. A euornithean known from a nearly complete skeleton. May have had a wading ecology.
-Kizylkumavis (1984): Late Cretaceous (Turonian) Uzbekistan. Known only from an arm bone.
-Kookne (2019): Late Cretaceous (Maastrichtian) Argentina. A possible neornithean known only from a partial shoulder bone.
-Kuszholia (1992): Late Cretaceous (Turonian) Uzbekistan. Known only from several fragmentary parts. May have been some other type of maniraptor.
-Lamarqueavis (2010): Late Cretaceous (Campanian–Maastrichtian) Argentina. Known from a partial shoulder bone.
-Largirostrornis (1997): Early Cretaceous (Aptian) China. A long-snouted enantiornithean. Possibly the same as Cathayornis.
-Lectavis (1993): Late Cretaceous (Maastrichtian) Argentina. A long-legged enantiornithean. Known only from hindlimbs. May have been the same as Enantiornis.
-Lenesornis (1986): Late Cretaceous (Turonian) Uzbekistan. Known only from hip vertebrae.
-Liaoningornis (1996): Early Cretaceous (Barremian–Aptian) China. An enantiornithean with a large breastbone and good perching ability. Once thought to have been a euornithean.
-Liaoxiornis (1999): Early Cretaceous (Aptian) China. An enantiornithean known from a juvenile with well-developed flight feathers. Very likely the juvenile of another known genus.
-Limenavis (2001): Late Cretaceous (Campanian–Maastrichtian) Argentina. Known from only a few fragmented forelimb bones.
-Linyiornis (2016): Early Cretaceous (Aptian) China. An enantiornithean known from a nearly complete skeleton with preserved ovarian follicles.
-Lonchodytes (1963): Late Cretaceous (Maastrichtian) U.S.A. A possible neornithean known from a partial foot.
-Longchengornis (1997): Early Cretaceous (Aptian) China. An enantiornithean known from a good specimen but not well studied. Possibly the same as Cathayornis.
-Longicrusavis (2010): Early Cretaceous (Barremian–Aptian) China. A close relative of Hongshanornis. Had long legs.
-Longipteryx (2001): Early Cretaceous (Aptian) China. A long-snouted enantiornithean well adapted for perching. May have fed on fish or insects, but known to have eaten fruit. Includes "Camptodontornis".
-Longirostravis (2004): Early Cretaceous (Barremian–Aptian) China. A long-snouted enantiornithean that may have been a mud prober or an arboreal insectivore.
-Longusunguis (2014): Early Cretaceous (Aptian) China. An enantiornithean with long, thin claws on its feet. Closely related to Bohaiornis. May have used continuous flapping flight.
-Maaqwi (2017): Late Cretaceous (Campanian) Canada. A diving euornithean. Possibly closely related to Vegavis, though also had similarities to procellariiforms.
-Magnusavis (2024): Late Cretaceous (Maastrichtian) U.S.A. A large enantiornithean.
-Martinavis (2007): Late Cretaceous (Campanian–Maastrichtian) Argentina, France, and U.S.A. An extremely widespread enantiornithean, but probably an overlumped taxon. May have been the same as Soroavisaurus. Five species have been named, M. cruzyensis, M. vincei, M. saltariensis, M. minor, and M. whetstonei.
-Meemannavis (2021): Early Cretaceous (Aptian) China. A euornithean known from a partial skull and some vertebrae.
-Mengciusornis (2019): Early Cretaceous (Aptian) China. A euornithean with teeth restricted to the tip of the upper jaw.
-Microenantiornis (2017): Early Cretaceous (Aptian) China. A small enantiornithean known from a complete specimen.
-Mirarce (2018): Late Cretaceous (Campanian) U.S.A. A large enantiornithean and the most completely known enantiornithean from North America. The first enantiornithean found with quill knobs. Probably closely related to Avisaurus.
-Mirusavis (2019): Early Cretaceous (Aptian) China. An enantiornithean whose holotype was found with medullary bone preserved across most of its skeleton.
-Monoenantiornis (2016): Early Cretaceous (Aptian) China. An enantiornithean known from a nearly complete subadult specimen.
-Musivavis (2022): Early Cretaceous (Aptian) China. Possibly closely related to Bohaiornis, but was much smaller.
-Mystiornis (2011): Early Cretaceous (Barremian–Aptian) Russia. A diving avialan, possibly an enantiornithean.
-Nanantius (1986): Early Cretaceous (Albian) Australia. The first (and so far only) named enantiornithean from Australia.
-Navaornis (2024): Late Cretaceous (Santonian–Campanian) Brazil. A toothless enantiornithean. A nearly complete skull is known.
-Neogaeornis (1929): Late Cretaceous (Maastrichtian) Chile. Some kind of seagoing neornithean, possibly closely related to Vegavis. One of the first Cretaceous avialans found in South America.
-Neuquenornis (1994): Late Cretaceous (Santonian) Argentina. An enantiornithean known from a partial skeleton. Eggs with embryos that may belong to this taxon are known.
-Noguerornis (1989): Early Cretaceous (Barremian) Spain. An enantiornithean known from a partial skeleton with preserved feathers.
-Omnivoropteryx (2002): Early Cretaceous (Aptian) China. Probably the same as Sapeornis.
-Orienantius (2018): Early Cretaceous (Hauterivian) China. An enantiornithean known from two very well-preserved specimens. May have used flap-gliding flight.
-Otogornis (1994): Early Cretaceous (Aptian?) China. An enantiornithean known from only forelimb bones with preserved feathers.
-Palintropus (1970): Late Cretaceous (Campanian–Maastrichtian) Canada and U.S.A. Once thought to have been an early galliform, but was probably closer to Apsaravis.
-Parabohaiornis (2014): Early Cretaceous (Aptian) China. Closely related to Bohaiornis.
-Parahesperornis (1984): Late Cretaceous (Coniacian–Santonian) U.S.A. A hesperornithean known from a nearly complete skeleton is known.
-Parahongshanornis (2011): Early Cretaceous (Aptian) China. A euornithean known from a nearly complete skeleton lacking a skull.
-Parapengornis (2015): Early Cretaceous (Aptian) China. An enantiornithean with fully pennaceous tail feathers. Has been suggested to have been specialized for trunk-climbing, but this has been disputed.
-Paraprotopteryx (2007): Early Cretaceous (Aptian) China. The first enantiornithean to be found preserved with two pairs of ribbon-shaped tail feathers.
-Parvavis (2014): Late Cretaceous (Turonian–Santonian) China. An enantiornithean known from a subadult specimen. One of the smallest known Mesozoic dinosaurs, even taking into account potential adult size.
-Pasquiaornis (1997): Late Cretaceous (Cenomanian) Canada. A hesperornithean possibly capable of flying and waddling. May have been a surface swimmer. Known from only leg bones and a skull bone. Two species have been named, P. hardiei and P. tankei.
-Patagopteryx (1992): Late Cretaceous (Santonian) Argentina. A flightless euornithean known from a nearly complete skeleton but not a complete skull.Had a forward-pointing first toe and short feet.
-Pengornis (2008): Early Cretaceous (Aptian) China. One of the largest Early Cretaceous enantiornitheans. Had small blunt teeth.
-Piscivoravis (2013): Early Cretaceous (Aptian) China. Known to have eaten fish. Had several unusual features for a euornithean, such as a large thumb claw.
-Piscivorenantiornis (2017): Early Cretaceous (Aptian) China. An enantiornithean thought to have eaten fish based on a mass of fish bones associated with the holotype, but this is uncertain given the disarticulated nature of the specimen and the similarity of the mass to fish coprolites.
-Platanavis (1992): Late Cretaceous (Turonian) Uzbekistan. Known only from hip vertebrae.
-Polarornis (2002): Late Cretaceous (Maastrichtian) Antarctica. Has been suggested to have been an early loon, but was possibly closely related to Vegavis. May have been flightless or nearly so. Possibly the same as Neogaeornis.
-Potamornis (2000): Late Cretaceous (Maastrichtian) U.S.A. A freshwater hesperornithean.
-Protopteryx (2000): Early Cretaceous (Hauterivian) China. One of the oldest known enantiornitheans, known from several nearly complete specimens. Had short, broad wings. Likely exhibited sequential molt of its wing feathers. May have used bounding or flap-gliding flight. Analysis of the melanosomes preserved in its feathers suggests that at least some of its wing feathers had black tips.
-Pterygornis (2015): Early Cretaceous (Aptian) China. A small enantiornithean. Grew rapidly for an enantiornithean, nearing adult size within a year.
-Qiliania (2011): Early Cretaceous (Aptian) China. An enantiornithean known from well-preserved hip and leg bones.
-Rapaxavis (2009): Early Cretaceous (Aptian) China. A long-snouted enantiornithean suggested to have been adapted for perching and mud probing, but the latter has been considered unlikely. Known from a complete skeleton. The feet may have been pamprodactyl.
-Sapeornis (2002): Early Cretaceous (Barremian–Aptian) China. A large omnivoropterygid with long foot feathers and very long wings. Its name stands for the Society of Avian Paleontology and Evolution. Known to have eaten seeds. The size and shape of its sclerotic ring suggest it may have been diurnal. Likely specialized for thermal soaring. Includes "Didactylornis" and "Shenshiornis".
-Sazavis (1989): Late Cretaceous (Turonian) Uzbekistan. Known only from a leg bone fragment.
-Schizooura (2012): Early Cretaceous (Aptian) China. A fork-tailed, toothless euornithean.
-Shangyang (2019): Early Cretaceous (Aptian) China. A small enantiornithean known from a nearly complete skeleton.
-Shanweiniao (2010): Early Cretaceous (Barremian–Aptian) China. A long-snouted enantiornithean. Its name means "fan-tailed bird" in Chinese, based on the fact that it was found with four to six ribbon-shaped tail feathers preserved instead of two.
-Shengjingornis (2012): Early Cretaceous (Aptian) China. An enantiornithean with a downcurved snout.
-Shenqiornis (2010): Early Cretaceous (Aptian) China. An enantiornithean found with a well-preserved skull. Had bulb-shaped teeth that indicate it may have fed on hard, durable food items.
-Shuilingornis (2024): Early Cretaceous (Aptian) China. Possibly closely related to Gansus.
-Similiyanornis (2020): Early Cretaceous (Aptian) China. Closely related to Yanornis, as its name suggests. Had a pair of unusually large teeth in its lower jaw.
-Sinornis (1992): Early Cretaceous (Aptian) China. An enantiornithean known from a nearly complete fossil.
-Songlingornis (1997): Early Cretaceous (Aptian) China. Probably a yanornithiform.
-Soroavisaurus (1993): Late Cretaceous (Maastrichtian) Argentina. Probably closely related to Avisaurus, which it was once considered a species of.
-Sulcavis (2013): Early Cretaceous (Aptian) China. A durophagous enantiornithean. Had a long hyoid for a non-neornithean theropod. May have used continuous flapping flight.
-Telmatornis (1870): Late Cretaceous–Paleocene? (Maastrichtian–Danian?) U.S.A. A possible neornithean known from limb bones.
-Teviornis (2002): Late Cretaceous (Maastrichtian) Mongolia. Probably a presbyornithid anseriform.
-Tianyuornis (2014): Early Cretaceous (Barremian–Aptian) China. A hongshanornithid known from a subadult specimen.
-Tingmiatornis (2016): Late Cretaceous (Turonian) Canada. A large diving euornithean.
-Torotix (1963): Late Cretaceous (Maastrichtian) U.S.A. A possible neornithean known from a partial arm bone. Although its name means "flamingo", it has been suggested to have been an aequornithean.
-Vegavis (2005): Late Cretaceous (Maastrichtian) Antarctica. An early diving neornithean, possibly an anseriform. One specimen has a preserved syrinx.
-Vescornis (2004): Early Cretaceous (Aptian) China. Was briefly described in 1999 as "Hebeiornis", but this name is not used by most researchers. May have been the adult of Jibeinia.
-Volgavis (1989): Late Cretaceous (Maastrichtian) Russia. Possibly a pelican-like marine euornithean.
-Vorona (1996): Late Cretaceous (Maastrichtian) Madagascar. Known only from limb bones.
-Wyleyia (1973): Early Cretaceous (Barremian) U.K. May have been some other type of maniraptor.
-Xiangornis (2012): Early Cretaceous (Aptian) China. An enantiornithean with a hand skeleton similar in structure to those of euornitheans.
-Xinghaiornis (2013): Early Cretaceous (Aptian) China. A toothless euornithean probably specialized for mud probing.
-Yangavis (2018): Early Cretaceous (Aptian) China. A confuciusornithiform with an unreduced claw on the second finger, unlike other confuciusornithiforms.
-Yanornis (2001): Early Cretaceous (Aptian) China. The front half of a specimen of this euornithean was once combined with the back half of a specimen of Microraptor to create the infamous "Archaeoraptor" hoax.
-Yatenavis (2022): Late Cretaceous (Maastrichtian) Argentina. A small enantiornithean known from a partial arm bone.
-Yixianornis (2001): Early Cretaceous (Aptian) China. Probably closely related to Yanornis. The size and shape of its sclerotic ring suggest it may have been diurnal.
-Yuanchuavis (2021): Early Cretaceous (Aptian) China. An enantiornithean with a fan-shaped array of tail feathers, superficially similar to that of euornitheans. The central pair of tail feathers was highly elongated. Analysis of the melanosomes preserved in its feathers suggests that its central pair of tail feathers may have been black, whereas its other tail feathers may have been gray.
-Yuanjiawaornis (2015): Early Cretaceous (Aptian) China. A large enantiornithean.
-Yumenornis (2013): Early Cretaceous (Aptian) China. A euornithean known from a complete forelimb and associated parts.
-Yungavolucris (1993): Late Cretaceous (Maastrichtian) Argentina. May have been an aquatic enantiornithean.
-Yuornis (2021): Late Cretaceous (Maastrichtian?) China. A toothless enantiornithean known from a partial skeleton, including a complete skull.
-Zhongjianornis (2010): Early Cretaceous (Aptian) China. Described as a toothless non-pygostylian avialan. May have actually been a euornithean.
-Zhongornis (2008): Early Cretaceous (Aptian) China. Known from a hatchling, perhaps a confuciusornithiform. Had a short tail but no pygostyle.
-Zhouornis (2013): Early Cretaceous (Aptian) China. A fairly large enantiornithean. May have used continuous flapping flight.
-Zhyraornis (1992): Late Cretaceous (Turonian) Uzbekistan. Known only from vertebrae.
Indeterminate/Miscellaneous maniraptors
-Alcmonavis (2019): Late Jurassic (Tithonian) Germany. An early avialan known from a forelimb.
-Ambopteryx (2019): Middle Jurassic (Callovian) China. A scansoriopterygid known to have had membranous wings. The holotype was preserved with gizzard stones.
-Anchiornis (2009): Late Jurassic (Oxfordian) China. May have been either a deinonychosaur or an avialan. Had long foot feathers and feathered toes. Known to have eaten fish and lizards. Analysis of the melanosomes preserved in its feathers indicates that it may have had a mostly black and dark gray body with an orange crown and black-and-white wing feathers.
-Archaeopteryx (1861): Late Jurassic (Tithonian) Germany. Known from several excellent fossils. Had long feathers on its legs, though not as extensive as those of Anchiornis or Microraptor. Analysis of its wing feathers show that they may have been black tipped. May have exhibited sequential molt of its wing feathers. The size and shape of its sclerotic ring suggest it may have been diurnal. May have been an avialan, deinonychosaur, or a paravian outside of both lineages. Includes "Wellnhoferia".
-Aurornis (2013): Late Jurassic (Oxfordian) China. Described as one of the earliest known avialans. May have been the same as Anchiornis.
-Balaur (2010): Late Cretaceous (Maastrichtian) Romania. A robust paravian with an enlarged first toe and just two functional fingers. May have been either a dromaeosaurid or an avialan.
-Bradycneme (1975): Late Cretaceous (Maastrichtian) Romania. Known only from a partial leg bone. Initially thought to have been an owl.
-Caihong (2018): Late Jurassic (Oxfordian) China. A paravian with small crests in front of its eyes. It had very long feathers on its wings and tail, and also had feathered toes. Analysis of the melanosomes preserved in its feathers suggests that it may have been mostly black, with iridescent plumage on its head and chest.
-Elopteryx (1913): Late Cretaceous (Maastrichtian) Romania. Has been suggested to have been a dromaeosaurid, but was probably a troodont or avialan.
-Eosinopteryx (2013): Late Jurassic (Oxfordian) China. An early paravian that appears to have lacked pennaceous tail feathers.
-Epidexipteryx (2008): Middle Jurassic (Callovian) China. A scansoriopterygid with a short snout, large protruding teeth, and a short tail. Was found with four long ribbon-shaped tail feathers preserved. Possibly an older individual of Scansoriopteryx.
-Euronychodon (1991): Late Cretaceous (Campanian–Maastrichtian) Portugal. Known only from teeth that may belong to deinonychosaurs.
-Fujianvenator (2023): Late Jurassic (Tithonian) China. An early paravian with unusually long lower legs and feet.
-Fukuipteryx (2019): Early Cretaceous (Aptian) Japan. May have been a non-pygostylian avialan despite its short tail.
-Fukuivenator (2016): Early Cretaceous (Aptian) Japan. Had heterodont dentition suggestive of omnivory, and a pygostyle that had evolved independently of pygostylians. Likely had a good sense of smell. May have been a therizinosaur, alvarezsaur, paravian, or some other type of coelurosaur.
-Imperobator (2019): Late Cretaceous (Maastrichtian) Antarctica. A large paravian known only from fragmentary feet.
-Jeholornis (2002): Early Cretaceous (Aptian) China. One of the largest Early Cretaceous avialans. Known to have eaten fruit and leaves. Had a first toe that was not fully reversed. Its tail feathers were frond-like and probably used in display. It also had modified tail coverts that formed a fan-shaped structure. The size and shape of its sclerotic ring suggest it may have been diurnal. Three species have been named, J. prima, J. curvipes, and J. palmapenis.
-Jixiangornis (2002): Early Cretaceous (Barremian–Aptian) China. Possibly the same as Jeholornis.
-Kakuru (1980): Early Cretaceous (Aptian) Australia. Known from only some limb bones that may belong to an oviraptorosaur. May have been an abelisauroid.
-Kompsornis (2020): Early Cretaceous (Aptian) China. Known from a nearly complete skeleton. Similar to Jeholornis.
-Migmanychion (2023): Early Cretaceous (Aptian) China. Known mostly from a partial forelimb, including a complete hand. Had some similarities to Fukuivenator.
-Neimengornis (2021): Early Cretaceous (Aptian) China. Known from a nearly complete skeleton with preserved tail feathers. Similar to Jeholornis.
-Nuthetes (1854): Early Cretaceous (Berriasian) U.K. May have been a dromaeosaurid.
-Ostromia (2017): Late Jurassic (Tithonian) Germany. The holotype was initially described as a species of Pterodactylus, and then a specimen of Archaeopteryx.
-Overoraptor (2020): Late Cretaceous (Cenomanian–Turonian) Argentina. Possibly closely related to Rahonavis.
-Paronychodon (1876): Late Cretaceous (Turonian–Campanian) U.S.A. and Uzbekistan. Known only from teeth that may belong to deinonychosaurs. Teeth from numerous locales have been referred to this taxon, but it is unclear whether they all represent the same genus.
-Pedopenna (2005): Middle Jurassic (Callovian) China. A paravian known only from a hindlimb with plumulaceous foot feathers preserved.
-Pneumatoraptor (2010): Late Cretaceous (Santonian) Hungary. A small paravian with very hollow bones.
-Rahonavis (1998): Late Cretaceous (Maastrichtian) Madagascar. A small, flying unenlagiine or avialan. Quill knobs have been identified on its ulna. Was initially named "Rahona", but that name had already been used for a lepidopteran.
-Richardoestesia (1990): Late Cretaceous (Turonian–Maastrichtian) Canada and Uzbekistan. Known only from jaws and teeth that may belong to deinonychosaurs. Teeth from numerous locales have been referred to this taxon, but it is unclear whether they all represent the same genus. May have been some other type of theropod.
-Scansoriopteryx (2002): Middle Jurassic (Callovian) China. Known from two juvenile specimens. Had an extremely long third finger and may have climbed trees. Includes "Epidendrosaurus". May have been an avialan.
-Serikornis (2017): Late Jurassic (Oxfordian) China. An early paravian known to have been covered in plumulaceous feathers down to its toes.
-Shenzhouraptor (2002): Early Cretaceous (Aptian) China. May have been the same as Jeholornis.
-Xiaotingia (2011): Late Jurassic (Oxfordian) China. Had blunt, bulbous teeth. May have been a deinonychosaur or avialan.
-Yandangornis (1999): Late Cretaceous (Campanian) China. A long-tailed, toothless avialan with long legs. May have been some other type of maniraptor.
-Yaverlandia (1971): Early Cretaceous (Barremian) U.K. Known only from the top of a skull. Initially thought to have been a pachycephalosaur.
-Yi (2015): Late Jurassic (Oxfordian) China. A scansoriopterygid known to have had an unusual styliform bone jutting from each forelimb and extensive patagia, forming membranous wings.
-Yixianosaurus (2003): Early Cretaceous (Barremian–Aptian) China. Known only from forelimb material with preserved feathers. Had very long hands. May have been a paravian.
Ilerdopteryx
ReplyDeletePraeornis
Thecocoelurus
Richardoestesia
Pectinodon
Thanks. I'm leaving out Praeornis as it's dubious, and provisionally keeping Pectinodon in Troodon. I intentionally left out Thecocoelurus as I remember reading on Tetrapod Zoology that Naish no longer considers it an oviraptorosaur or therizinosaur, so I'm waiting for further info. I didn't include Richardoestesia because I based much of the phylogenetics on Holtz's winter 2010-2011 dinosaur genus list, where it is only categorized as a basal coelurosaur. I might consider adding Ilerdopteryx.
ReplyDeleteOn second thought, I'll hold off Ilerdopteryx as well, it being non diagnostic.
ReplyDeleteGreeting Albertonykus
ReplyDeleteParascaniornis stensioei
Polarornis
Hebeiornis (Vescornis)
Anchiornis
D G Rexisto
Greetings. Thanks for the feedback. I'm keeping Parascaniornis in Baptornis, and Hebeiornis, it appears, isn't "adequately described". Anchiornis is already there. I'll add Polarornis.
ReplyDeleteGood job! You deserve points for holding off on Chatterjee's Protoavis!
ReplyDeleteRelated note: How are we sure that Yaverlandia is a maniraptor and not some other theropod? All we've got is a skull roof!
"I intentionally left out Thecocoelurus as I remember reading on Tetrapod Zoology that Naish no longer considers it an oviraptorosaur or therizinosaur, so I'm waiting for further info."
So? He's just one person. Not that he's unreliable in any way, but he's still just one person.
Not to mention, what you said here about Eshanosaurus was cool. I didn't know that.
Definitely not touching "Protoavis"! XD
ReplyDeleteYaverlandia as a maniraptor is derived from this analysis: http://jgs.geoscienceworld.org/cgi/content/abstract/165/3/613
On Thecocoelurus, it's mostly that Naish was the one who initially identified it as a maniraptor to begin with. I think he's said that he now finds that it's an abelisauroid, but wait for the paper, I guess.
"Reproductive organs may be preserved in one individual."
ReplyDeleteWhich gender are we talking about, by the way?
While we're at it, could you please move the scansoriopterygids to the "Indeterminate maniraptors" list? They're still controversial.
In my previous comment about Yaverlandia, I meant to ask how the study came to the conclusion that it was a maniraptor and not another type of theropod.
I forgot to mention: Is Eshanosaurus still that old? I recall reading in Dr. Holtz's syllabus that it may be as young as the Early Cretaceous.
ReplyDeleteThe reproductive organs are those of a male.
ReplyDeleteAlthough I personally expect (or at least wouldn't be surprised if) scansoriopterygids to turn out to be non avialians, most analyses that have included them have resulted in them being avialians, so they will remain there for now. The only recent exception I'm aware of is the Samrukia paper, though I could easily be wrong.
I don't have the Yaverlandia as maniraptor paper unfortunately, so I don't know which characters resulted in it being a maniraptor.
The exact age of Eshanosaurus is indeed controversial.
Struthio
ReplyDeleteCorvus
Pica
Haliaeetus
Many other modern birds. :P
From the introductory paragraph:
ReplyDelete"I hope to include the Cenozoic taxa someday, but for now I'm focusing on the Mesozoic ones."
Is that because there are too many to list? :P
ReplyDeleteYes. That, and I don't know of any handy databases that have them all compiled (particularly extinct taxa).
ReplyDeleteShould Yaverlandia be moved? I read on TetZoo that Naish specifically found it to be troodontid...
ReplyDeleteSpinosegnosaurus77/SpongeBobFossilPants
Nice list! Very handy. I don't have the paper either but here's what TTD says on Yaverlandia...
ReplyDelete"Naish believes it is a theropod, based on bilobed cerebral concavity; narrow olfactory tract; ventrally concave orbital margins; small, closely appressed olfactory bulbs. Indeed, the deep cerebral fossae and posteriorly expanded cerebrum suggest a coelurosaurian identity, while the narrow olfactory lobes are only known in maniraptoriforms. Among maniraptoriforms, Yaverlandia is unlike ornithomimids and dromaeosaurids in having ossified orbitosphenoids."
Hey, if you update this, could you add who described each maniraptoran?
ReplyDeleteThat might be a good idea!
DeleteA pronunciation guide would also be handy, as well as the inclusion of "Ingenia".
DeleteSpinosegnosaurus77/SpongeBobFossilPants
Not including non-validly named taxa.
DeleteAbout Eshanosaurus... I got in touch with Holtz concerning the genus list. He had this to say: "I know the paper [suggesting it's a therizinosaur], and am not impressed."
ReplyDeleteSpinosegnosaurus77/SpongeBobFossilPants (the previous comment was also mine, but I couldn't sign it for some reason)
Yes, he told me something similar when I asked him about it last year.
DeleteIf I ask him why he's unimpressed, will you remove it?
DeleteSpongeBobFossilPants/Spinosegnosaurus77
Probably not (I'd rather wait for something to come up in the technical lit before doing that), though I might note that there is doubt on the study's validity.
DeleteAbout Chirostenotes.. How do we know the feathers didn't run the length of the shorter two fingers, leaving the longer one bare?
ReplyDeleteSpinosegnosaurus77/SpongeBobFossilPants
Remiges attach to the second finger, not any other (except maybe in scansors, but they're weirdos so they get a pass), even if the other fingers are also feathered (as is usually the case, though possibly not for oviraptorosaurs).
DeleteI wasn't suggesting they didn't attach to the second finger, I suggested that they were shorter than usual.
DeleteSpinosegnosaurus77/SpongeBobFossilPants
That's entirely possible, but unless they were as short as penguin wing feathers or something I don't think they would have been conducive to probing.
DeleteShould we move unenlagiines, microraptorines & troodontids to Indeterminate Maniraptors?
ReplyDeleteSpinosegnosaurus77/SpongeBobFossilPants
Although all of those have the potential to jump around the tree, I don't think that's necessary for now.
DeleteWill you at least note that they may jump around the tree?
DeleteSpinosegnosaurus77/SpongeBobFossilPants
If it's recovered seriously by more analyses.
DeleteYou should really keep a Thescelosaurus!-like updates list.
ReplyDeleteSpinosegnosaurus77/SpongeBobFossilPants
Wasn't Epidendrosaurus named before Scansoriopteryx?
ReplyDelete-Ornitholestes
Sort of but not really.
DeleteAnd wasn't Shenzouraptor named before Jeholornis?
DeleteYeah... maybe. It's hard to tell because the two names were published within days of one another. I'm really just following the fact that Jeholornis is currently the favored name in technical literature, but the case has certainly been made for "Shenzhouraptor".
DeleteBy any chance, is there any real reason to ignore Mortimer's arguments (and those on Hebeiornis being the correct name for Vescornis)?
DeleteSpinosegnosaurus77/SpongeBobFossilPants
As far as I'm aware... no, probably not.
DeleteMickey Mortimer considers Inosaurus a therizinosaur; will you add it?
ReplyDeleteSpinosegnosaurus77/SpongeBobFossilPants
Maybe if it turns out to be less dubious...
DeleteIt might be a good idea to mention "Macrophalangia" in the comments for Chirostenotes.
ReplyDeleteSpinosegnosaurus77/SpongeBobFossilPants
Mickey Mortimer considers Kuszholia an oviraptorosaur; care to mention that?
ReplyDeleteSpinosegnosaurus77/SpongeBobFossilPants
Shouldn't you remove the part about two species of Sinornithosaurus?
ReplyDeleteWhich sources synonymize Achillesaurus with Alvarezsaurus (other than Paul's 2010 field guide)?
Spinosegnosaurus77/SpongeBobFossilPants
Waiting a bit to see if that conclusion is well supported.
DeleteAchillesaurus being Alvarezsaurus is suggested in the description of Alnashetri.
Khaan as a juvenile Citipati? Don't tell me you're using Paul's 2010 field guide as a reference!
ReplyDeleteSpinosegnosaurus77/SpongeBobFossilPants
Only because the possibility is backed up by Martyniuk's guide, if that counts for anything.
DeleteYou do realize that Macrophalangia was named before Caenagnathus, right?
ReplyDeleteAlso, you forgot Wulatelong.
SpongeBobFossilPants/Spinosegnosaurus77
I need to reread the Leptorhynchos paper.
DeleteAnd no, I did not, I just haven't added it.
Here we go: "No other jaws of Caenagnathus have been identified. CMN 2367, an articulated foot described as “Macrophalangia canadensis,” is usually assumed to be synonymous with Chirostenotes pergracilis, but it has a larger and more robust tarsometatarsus than Chirostenotes, which raises the possibility that it represents the same species as Caenagnathus. “Macrophalangia canadensis” Sternberg 1932 would have priority over Caenagnathus collinsi Sternberg 1940, but at present it is difficult to confidently refer the pes and jaw to the same species. Therefore, for now we prefer to retain the familiar Caenagnathus over the little-used “Macrophalangia.”"
DeleteI'll edit the list to clarify this.
Kakuru?
ReplyDeleteIt's there. ;)
DeleteAh, true... anyway, excellent list, I think it can be very useful to those who are beginners.
DeleteI would like to know what you think of a my speculation: Bauxitornis and tarsometatarsus of Balaur share certain traits in common. And if these common traits were synapomorphies? A possible very endemic clade of Avialae just outside Pygostylia including terrestrial and dwarfic taxa as Balaur, Bauxitornis and perhaps other doubts Hateg's maniraptorians, as Bradycneme, Elopteryx, Heptasteornis (these three lack in your list) and Pneumatoraptor: The mysterious Bradycnemidae!
Very interesting speculation; it wouldn't greatly surprise me if that were true. Better specimens are needed to properly evaluate it, naturally.
DeleteThose three are in the list, though I've put them under more specific categories than perhaps they should be.
Nqwebasaurus came out as an alvarezsaur in Lee et al. (2014); will you add it?
ReplyDeleteSpongeBobFossilPants/Spinosegnosaurus77
I'll wait until it starts showing up more consistently as one.
DeleteWhere has synonymy of Incisivosaurus with Protarchaeopteryx been suggested? Mickey Mortimer said in his review of Martyniuk's book that the idea was completely new to him.
ReplyDeleteSpinosegmosaurus77/SpongeBobFossilPants
The original suggestion was probably that of Senter et al. (2004) "Systematics and evolution of Dromaeosauridae (Dinosauria, Theropoda)".
DeleteAren't "Palaeopteryx" and "Proornis" nomina nuda?
ReplyDeleteAlso, do you use [] or <> for italics?
Spinosegnosaurus77/SpongeBobFossilPants
That appears to be the case. I'll exclude them for now.
Delete<> for italics (and other formatting tags).
Which analyses (other than the original description) placed Jinfengopteryx as an avialan (without other troodontids following)?
ReplyDeleteHas anyone seriously followed Agnolin & Novas (2013) in supporting non-dromaeosaurid Mahakala?
Wasn't there a recent paper that had Richardoestesia as a dromaeosaurid?
What led you to remove Unquillosaurus? Was there a followup to Carrano et al. (2012) that I missed?
Spinosegnosaurus77/SpongeBobFossilPants
Non-troodont Jinfengopteryx was also found by Foth et al. (2014) describing the eleventh Archaeopteryx specimen.
DeleteI'm stumped on Mahakala... it doesn't appear to have been any of the usual suspects. The Theropod Database does say that it may be a basal paravian, but doesn't cite a source for it. I may have simply been mistaken in moving it out.
Richardoestesia I could go either way on.
With Unquillosaurus, I probably eventually decided on Carrano et al. (2012) being sufficient to justify removal until further notice. (It was a while ago.) I was likely also influenced by other sites (mostly Thescelosaurus!) that started including it among carnosaurs instead.
I thought Jinfengopteryx was found to be a basal paravian, not an avialan.
DeleteSpinosegnosaurus77/SpongeBobFossilPants
I misread your first comment. Yes, you're correct on that count.
DeleteSurely "Dalianraptor" isn't an avebrevicaudan… is it?
ReplyDeleteSpinosegnosaurus77/SpongeBobFossilPants
Forgot I still had it up there! Correct, it wouldn't be... if it's real.
DeleteDidn't Kurochkin (2001) show that Holbotia was distinct from Ambiortus?
ReplyDeleteFor that matter, isn't Holbotia a nomen nudum?
Spinosegnosaurus77/SpongeBobFossilPants
Yep.
DeleteWhich version of Pelecaniformes are you using?
ReplyDeleteSpinosegnosaurus77/SpongeBobFossilPants
The traditional, more inclusive sense (including cormorants). I'll likely change that... at some point.
DeleteAre you up for adding Praeornis now that we’re reasonably sure it’s avian?
ReplyDeleteI probably won't, given that it's unlikely that we would be able to confirm it as belonging to a specific species even if we hypothetically found a more complete specimen. If we do manage to do that someday though, I'll certainly add it.
DeleteI've noticed in these lists you've included some dubious taxa (e.g. Nuthetes) while excluding others (e.g. Ilerdopteryx). I'm curious how you've decided which ones to include and which ones to leave out.
ReplyDeleteI haven't been very consistent about it, I'll admit, and it's something I should reevaluate when I have the time. Much of it is a function of whichever sources I was consulting when I originally made the list.
DeleteThat being said, I do draw a hard line against the inclusion of feather taxa like Ilerdopteryx, given how difficult it would be to confirm their association with skeleton-based genera.
What’s your justification for including Haplocheirus but not Tugulusaurus or Aorun?
ReplyDeletePrimarily the fact that alvarezsaurian affinities for Haplocheirus have been supported by analyses of more than one dataset (namely TWiG and Cau's datasets).
Delete